ISTANBUL ANKARA - Turkish President Recep Tayyip Erdogan has claimed that former Egyptian President Mohammed Morsi didn’t die of natural causes, but that he was killed.

During a speech in Istanbul, Erdogan cited as evidence that the deposed Egyptian president allegedly “flailed” in a Cairo courtroom for 20 minutes Monday and nobody assisted him.

The Turkish leader said Wednesday: “Unfortunately, Mohammed Morsi was on the ground of the courtroom flailing for 20 minutes. No official there intervened. Morsi did not (die) naturally, he was killed. Erdogan said Turkey would do everything in its power to ensure Egypt faces trial in Morsi’s death. He also called on the Islamic Cooperation Organization to “take the necessary action” over Morsi’s death.”

A previous version of this story was corrected to show that the speech was made Wednesday, not Thursday.

Former Egyptian President Mohammed Morsi has been buried hours after he collapsed in court and died on Monday. His lawyer told the AFP news agency the former leader had been buried in eastern Cairo early on Tuesday morning with his family present. Morsi, who was 67, had been in custody since his removal from office in 2013. Human rights groups, who had criticised the conditions in which he was kept, have called for an impartial investigation into his death.

His family and activists had repeatedly raised concerns about his health and the amount of time he was kept in solitary confinement, away from visits by lawyers and family. His son, Abdullah Mohamed Morsi, told Reuters on Monday that Egyptian authorities had denied a family request for a public funeral in his hometown.

A police presence was seen at the cemetery where Morsi was buried in Cairo. A top figure in the now-banned Islamist movement, the Muslim Brotherhood, Morsi became the country’s first democratically elected leader in 2012.

He was ousted and detained in a military coup a year later following mass protests against his rule.

Abdul Fattah al-Sisi, a former army chief, has been in power since 2014.

After Morsi’s removal, the authorities launched a crackdown on his supporters and other dissent, leading to tens of thousands of arrests. The Muslim Brotherhood and Turkish President Recep Tayyip Erdogan, a close ally of Morsi, are among those who have blamed the Egyptian leadership for Morsi’s death.

What happened in court?

Morsi was appearing in Cairo on Monday on charges of espionage related to suspected contacts with the Palestinian militant Islamist group Hamas.

Officials say he had asked to address the jury, and spoke for about five minutes from a soundproof glass cage where he was being held with other defendants.

Minutes later, he apparently fainted during a break in proceedings.

“He was transferred to a hospital where he was pronounced dead,” a statement by Egypt’s public prosecutor said. Officials said a forensic report had been ordered into his death, and insisted no new, visible injuries were found on his body.

State television had earlier described the cause of death as a heart attack.

Morsi was already facing decades in jail after being sentenced in three other trials. He had previously been given a death sentence, which was later overturned.

What has the reaction been?

Sarah Leah Whitson, executive director of Human Rights Watch’s Middle East and North Africa division, tweeted that his death was “terrible but entirely predictable”. “Former President Morsi’s death followed years of government mistreatment, prolonged solitary confinement, inadequate medical care, and deprivation of family visits and access to lawyers,” Ms Whitston said in a later statement.

The group called on the UN to start an investigation into what it described as “ongoing gross violations of human rights in Egypt, including widespread ill-treatment in prisons”. Amnesty International has also called for an impartial inquiry.

The rights group says Morsi was only allowed three family visits in almost six years of solitary confinement, and was denied access to his lawyers or a doctor.

The Muslim Brotherhood’s political arm, the Freedom and Justice party, described Morsi’s death as an “assassination”. Turkish President Erdogan blamed Egypt’s “tyrants” for his death and described him as a martyr.

The emir of Qatar, Sheikh Tamim bin Hamad Al Thani, another ally of the late leader, expressed his “deep sorrow” at the death. British MP Crispin Blunt, who led a panel of politicians who warned in 2018 about the conditions of Morsi’s confinement, has called for a “reputable independent international investigation”.